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Later, AUC was used to measure the size and salt-dependent assembly state of histones and to identify and characterize the core histone octamer, a complex of eight histone proteins (two each of H2A, H2B, H3, and H4), with derived results being essential for developing the nucleosome model of chromatin organization (Kornberg, 1974

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Astragaloside IV (AST-IV) (Fig