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PMID 12861015

The thread on tests for PA and B12 deficiency mentions other tests that can help to diagnose PA eg gastrin test and others (link in my other reply) Forum Support in reply to When you were tested for coeliac disease

Written By Martin Weaver, MD Dr

Ferreira et al., 2021

These four genes encode four different proteins that exhibit tissue-specific patterns of expression and exert distinct roles in cell differentiation and growth control

The third term is the loss of cysteine in the reaction catalyzed by GCS that makes glutamyl-cysteine and the fourth term represents the loss of cysteine to other pathways (for example to sulfate and taurine)